Betty has a hard time telling Daniel important news about her future; Amanda copes with the death of her dog; Marc wonders if he will ever have a successful relationship.

As Hilda’s wedding day approaches, Betty asks Daniel to be her plus-one, Ignacio invites Elena, and Amanda is set on fixing up Marc with her new client, soap opera star Spencer Cannon (Bryan Batt). Meanwhile, Justin weighs whether his own plus-one could be Austin, Betty considers unexpected offers in her personal and professional life, and Wilhelmina finds a way to fight the Meade empire by using Tyler as her pawn.

Betty brings Hilda on an assignment to cover London Fashion Week; Willie joins AA to get closer to Tyler and to double-cross Claire and Daniel; Bobby wonders why Justin is spending time with Austin.

After being knocked unconscious at a photo shoot, Betty has a dream in which her orthodontist (Kathy Najimy) shows her how her life would have been had she never worn braces.

Betty is pressured into dating an annoying firefighter (Rich Sommer); Amanda persuades Claire to hire Tyler; Hilda meets Bobby’s parents.

Daniel helps Betty to cover a low-priority show. Betty discovers a new designer, Marisa, and decides to promote her however she faces stiff competition from Marc. Amanda struggles to persuade Marc or Betty to vouch for her to Wilhemina about her clothing line. Tyler tries to find Claire in New York.

Betty needs to get a diamond-encrusted bra across town, and she has three ways of getting there

Betty tries to be neighborly by planning a karaoke party in her apartment building, but before the big day, her building is robbed, and it’s partly her fault. Marc is distraught since he lost something irreplaceable in the theft. Meanwhile, Daniel and Wilhelmina are back to being at each other’s throats, so Amanda tricks them into attending couples counseling with Dr. Sparks (Frank Whaley), and Ignacio refuses to accept Bobby in...

Betty recalls everything that has made up her worst week in history, and also has job issues caused by taking up a new hobby. Hilda is treated to a week of luxury by Bobby, and Daniel is becoming suspicious of Mark’s actions.

Betty wants to spend some time apart from Matt, so finds him something else to keep him occupied. Meanwhile, Hilda has a tough decision to make regarding her baby, and Mark’s fling looks to be heading towards something more serious.

Daniel hands over the power to the Community of the Phoenix to run Mode. Betty steps in to help Daniel with Bennet. She comes up with an cunning plan with the help of Claire. Wilhemina goes out in public wearing sweats creating mayhem with the media trying to get a scoop.

Marc encourages Justin to befriend the cheerleaders at school; Bennett and Natalie lure Daniel deeper into the Community of the Phoenix; Wilhelmina tries to find money for Nico’s blackmail payment.

At Wilhemina’s fund-raiser for Tibetan orphans Matt asks someone to be his partner. Daniel hires Natalie as his assistant, making Betty feel unneeded. Daniel finds out that Claire has a secret.

Betty has Marc as be her temporary assistant thanks to Wilhemina. Betty ends up covering the worst jobs in New York as an assignment but she needs to dress up as a hot dog and perform a Bollywood style dance. Marc has insecurities about how his idol at Vogue magazine will percieve him. Daniel is targeted by the leader of the Community.

Betty tries to get a scoop with Gucci’s new designer, Evan York, through Hilda’s customer, Sammy. Marc tries to stop her by getting Amanda and Matt to stage a showdown at Ignacio’s restaurant. Daniel goes to a bereavement group and meets Natalie who shows him an interesting night. Wilhelmina flies to Bermuda to try and find Connor.

Betty gets to run her own Mode photo shoot at the United Nations to promote their anti-malaria initiative, “Nothing but Nets,” but her big opportunity becomes a baptism by fire when Daniel’s neediness takes its toll, and Matt openly demoralizes her. Meanwhile, Amanda does some soul-searching when she bonds with Daniel’s new temp, Helen, a career receptionist as fabulous as Amanda is. Wili makes it her personal...

Betty starts her new job as associate features editor and deals with resentful co-workers; bullies prey on Justin; Wilhelmina harbors a secret; Daniel’s distress takes its toll, while Betty tries to run her own Mode photo shoot.